Steven D. Shema

Research Associate

Steven D. Shema

Grótti ehf
Grundarstíg 4
101 Reykjavík

The image below is a word cloud that is descriptive of Steve’s research activity.


M.A. University of Iceland
B.A. New York University


Recent Professional Experience

2016–Present, Managing Director, Grótti ehf
2009–2016, Independent Research Associate
2008–2013, Research Technician, Bigelow Laboratory for Ocean Sciences

Biographical Statement

Coming from a background in the performing arts and Old Norse literature, I have been engaged in marine science since 2008, primarily focusing on sensory ecology and the effects of climate change.

Research Interests or Outline of Research

Zooplankton behavior and ecology
Effects of climate change on plankton
Wild stock monitoring of wrasse, lumpfish and European eels

Current Projects

Effect of ocean acidification on Calanus spp. and other copepods
Fine-scale interactions in the plankton
In situ swimming and orientation ability of larval fishes and invertebrates
Population dynamics of wrasses along the Norwegian coast


Fields DM, Durif CMF, Bjelland RM, Shema SD, Skiftesvik AB, Browman HI (2011). Grazing Rates of Calanus finmarchicus on Thalassiosira weissflogii Cultured under Different Levels of Ultraviolet Radiation. Plos One, 6(10).
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Fields DM, Shema SD, Browman HI, Browne TQ, Skiftesvik AB (2012). Light Primes the Escape Response of the Calanoid Copepod, Calanus finmarchicus. Plos One, 7(6).
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Fukunishi Y, Browman HI, Durif CMF, Bjelland RM, Shema SD, Fields DM, Skiftesvik AB (2013). Sub-lethal exposure to ultraviolet radiation reduces prey consumption by Atlantic cod larvae (Gadus morhua). Marine Biology, 160(10), 2591-6.
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Nuester J, Shema S, Vermont A, Fields DM, Twining BS (2014). The regeneration of highly bioavailable iron by meso- and microzooplankton. Limnology and Oceanography, 59(4), 1399–1409.
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Fields DM, Runge JA, Thompson C, Shema SD, Bjelland RM, Durif CMF, Skiftesvik, AB, Browman HI (2015). Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp.: effects on grazing, respiration, fecundity, and fecal pellet production. Journal of Plankton Research 37(1) 211–20.
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Durif CMF, Fields DM, Browman HI, Shema SD, Enoae JR, Skiftesvik AB, Bjelland RM, Sommaruga R, Arts MT (2015). UV radiation changes algal stoichiometry but does not have cascading effects on a marine food chain. Journal of Plankton Research 37 (6), 1120–36.
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Runge JA, Fields DM, Thompson CRS, Shema SD, Bjelland RM, Durif CMF, Skiftesvik AB, Browman HI (2016). End of the century CO2 concentrations do not have a negative effect on vital rates of Calanus finmarchicus, an ecologically critical planktonic species in North Atlantic ecosystems. ICES Journal of Marine Science 73(3): 937-950.
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Bailey A, Thor P, Browman H, Fields, DM, Runge, J, Vermont A, Bjelland R, Thompson C, Shema S, Durif C, Hop H. (2017). Early life stages of the Arctic copepod Calanus glacialis are unaffected by increased seawater pCO2. ICES Journal of Marine Science (in press).
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Bailey, A., P. de Wit, P. Thor, H.I. Browman, R.M. Bjelland, S. Shema, D.M. Fields, J.A. Runge, C. Thompson & H. Hop. 2017. Regulation of gene expression underpins tolerance of the Arctic copepod Calanus glacialis to increased pCO2. Ecology and Evolution (in press).

Last updated: 20 March 2017