Steven D. Shema

Research Associate

Grótti ehf
Grundarstíg 4
101 Reykjavík
Iceland
Email: steven.shema@gmail.com

Education

M.A. University of Iceland
B.A. New York University

Recent Professional Experience

2016–Present, Managing Director, Grótti ehf
2009–2016, Independent Research Associate
2008–2013, Research Technician, Bigelow Laboratory for Ocean Sciences

Biographical Statement

Coming from a background in the performing arts and Old Norse literature, I have been engaged in marine science since 2008, primarily focusing on sensory ecology and the effects of climate change.

Research Interests

  • Zooplankton behavior and ecology
  • Effects of climate change on plankton
  • Monitoring and population dynamics of wild stocks of wrasse, lumpfish and European eel

Active projects

  • Effect of ocean acidification on Calanus spp. and other copepods (P.I. Howard Browman)
  • Fine-scale interactions in the plankton (P.I. Howard Browman)
  • In situ swimming and orientation ability of larval fishes and invertebrates (P.I. Howard Browman)
  • Population dynamics of wrasses along the Norwegian coast (P.I. Anne Berit Skiftesvik)

Publications

Regulation of gene expression underpins tolerance of the Arctic copepod Calanus glacialis to increased pCO2.

Bailey, A., P. de Wit, P. Thor, H.I. Browman, R.M. Bjelland, S. Shema, D.M. Fields, J.A. Runge, C. Thompson & H. Hop. 2017. Regulation of gene expression underpins tolerance of the Arctic copepod Calanus glacialis to increased pCO2. Ecology and Evolution (in press).

Glass eels (Anguilla anguilla) have a magnetic compass linked to the tidal cycle.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik & H.I. Browman. 2017. Glass eels (Anguilla anguilla) have a magnetic compass linked to the tidal cycle. Science Advances 3: e1602007, DOI: 10.1126/sciadv.1602007.
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The early life stages of the Arctic copepod Calanus glacialis are unaffected by increased seawater pCO2.

Bailey, A., P. Thor, H.I. Browman, D.M. Fields, J.A. Runge, A. Vermont, R. Bjelland, C. Thompson, S. Shema, C.M.F. Durif & H. Hop. 2017. The early life stages of the Arctic copepod Calanus glacialis are unaffected by increased seawater pCO2. ICES Journal of Marine Science.
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End of the century CO2 concentrations do not have a negative effect on vital rates of Calanus finmarchicus, an ecologically critical planktonic species in North Atlantic ecosystems.

Runge, J.A., D.M. Fields, C. Thompson, S. Shema, R.M. Bjelland, C.M.F. Durif, A.B. Skiftesvik & H.I. BROWMAN. 2016. End of the century CO2 concentrations do not have a negativeeffect on vital rates of Calanus finmarchicus, an ecologically criticalplanktonic species in North Atlantic ecosystems.ICES Journal of Marine Science 73: 937-950.
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Early life stages of the Arctic copepod Calanus glacialis are unaffected by increased seawater pCO2.

Bailey A, Thor P, Browman H, Fields, DM, Runge, J, Vermont A, Bjelland R, Thompson C, Shema S, Durif C, Hop H. (2016). Early life stages of the Arctic copepod Calanus glacialis are unaffected by increased seawater pCO2. ICES Journal of Marine Science: Journal du Conseil, fsw066.

End of the century CO2 concentrations do not have a negative effect on vital rates of Calanus finmarchicus, an ecologically critical planktonic species in North Atlantic ecosystems.

Runge JA, Fields DM, Thompson CRS, Shema SD, Bjelland RM, Durif CMF, Skiftesvik AB, Browman HI (2016). End of the century CO2 concentrations do not have a negative effect on vital rates of Calanus finmarchicus, an ecologically critical planktonic species in North Atlantic ecosystems. ICES Journal of Marine Science 73(3): 937-950.
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UV radiation changes algal stoichiometry but does not have cascading effects on a marine food chain.

Durif CMF, Fields DM, Browman HI, Shema SD, Enoae JR, Skiftesvik AB, Bjelland RM, Sommaruga R, Arts MT (2015). UV radiation changes algal stoichiometry but does not have cascading effects on a marine food chain. Journal of Plankton Research 37 (6), 1120–36.
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Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp.: effects on grazing, respiration, fecundity, and fecal pellet production.

Fields, D.M., J.A. Runge, C. Thompson, S.D. Shema, R.M. Bjelland, C.M.F. Durif, A.B. SKIFTESVIK & H.I. Browman. 2015. Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp.: effects on grazing, respiration, fecundity, and fecal pellet production. Journal of Plankton Research 37: 211-220.
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Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp.: effects on grazing, respiration, fecundity, and fecal pellet production.

Fields DM, Runge JA, Thompson C, Shema SD, Bjelland RM, Durif CMF, Skiftesvik, AB, Browman HI (2015). Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp.: effects on grazing, respiration, fecundity, and fecal pellet production. Journal of Plankton Research 37(1) 211–20.
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Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp: effects on grazing, respiration, fecundity and fecal pellet production.

Fields, D. M., Runge, J. A., Thompson, C., Shema, S. D., Bjelland, R. M., Durif, C. M. F., Skiftesvik, AB & Browman, H. I. (2015). Infection of the planktonic copepod Calanus finmarchicus by the parasitic dinoflagellate, Blastodinium spp: effects on grazing, respiration, fecundity and fecal pellet production. Journal of Plankton Research, 37(1), 211-220.
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The regeneration of highly bioavailable iron by meso- and microzooplankton.

Nuester J, Shema S, Vermont A, Fields DM, Twining BS (2014). The regeneration of highly bioavailable iron by meso- and microzooplankton. Limnology and Oceanography, 59(4), 1399–1409.
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Sub-lethal exposure to ultraviolet radiation reduces prey consumption by Atlantic cod larvae (Gadus morhua).

Fukunishi Y, Browman HI, Durif CMF, Bjelland RM, Shema SD, Fields DM, Skiftesvik AB (2013). Sub-lethal exposure to ultraviolet radiation reduces prey consumption by Atlantic cod larvae (Gadus morhua). Marine Biology, 160(10), 2591-6.
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Effects of ocean acidification on growth and development of the planktonic copepod,Calanusfinmarchicus.

Runge, J.A., C.Thompson, R.M. Bjelland, H.I. BROWMAN, C.M.F. Durif, D.M. Fields, S. Shema & A.B. Skiftesvik. 2013.Effects of ocean acidification on growth and development of the planktonic copepod,Calanusfinmarchicus. Poster presented at the U.S. National Science Foundation’s 2ndU.S. Ocean Acidification Principal Investigator’s Meeting. Washington, D.C., USA, 18-20 September 2013.
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Light Primes the Escape Response of the Calanoid Copepod, Calanus finmarchicus.

Fields DM, Shema SD, Browman HI, Browne TQ, Skiftesvik AB (2012). Light Primes the Escape Response of the Calanoid Copepod, Calanus finmarchicus. Plos One, 7(6).
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Grazing Rates of Calanus finmarchicus on Thalassiosira weissflogii Cultured under Different Levels of Ultraviolet Radiation.

Fields,D.M., C.M.F. Durif, R.M. Bjelland, S.D. Shema, A.B. Skiftesvik & H.I. BROWMAN. 2011. Grazing rates of Calanus finmarchicus on Thalassiosira weissflogii cultured under different levels of ultraviolet radiation.PLoS ONE 6(19) e26333.
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Grazing rates ofCalanus finmarchicuson diatoms cultured under different levels of ultraviolet radiation.

Fields,D.M., C.M.F. Durif, R.M. Bjelland, S.D. Shema, A.B. SKIFTESVIK & H.I. Browman. 2011. Grazing rates of Calanus finmarchicus on diatoms cultured under different levels of ultraviolet radiation. PLoS ONE6(10) e26333.
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Interaction between temperature and pH on growth and respiration of the planktonic copepod, Calanus finmarchicus. ICES Journal of Marine Science.

Runge, J.A., David Fields, Cameron Thompson, Caroline Durif, Steven Shema, Reidun Bjelland, Anne-Berit Skiftesvik and Howard I. Browman. Interaction between temperature and pH on growth and respiration of the planktonic copepod, Calanus finmarchicus. ICES Journal of Marine Science.

Vital rates of an ecologically critical planktonic species in North Atlantic ecosystems, Calanus finmarchicus, are unaffected by high CO2 but are affected by temperature.

Runge, J.A., D.M. Fields, C. Thompson, S. Shema, R.M. Bjelland, C.M.F. Durif, A.B. Skiftesvik, M. Arts, A. Mount, V. Chan & H.I. Browman. Vital rates of an ecologically critical planktonic species in North Atlantic ecosystems, Calanus finmarchicus, are unaffected by high CO2 but are affected by temperature.

Effect of ocean acidification on the respiration rates of Calanus finmarchicus.

Fields, D.M., H.I. Browman, A.B. SKIFTESVIK & S. Shema. Effect of ocean acidification on the respiration rates of Calanus finmarchicus.

Ontogeny of orientation in Atlantic haddock (Melanogrammus aeglefinus) larvae measured in situ.

Paris, C.B., H.I. Browman, A.B. Skiftesvik, S. Shema, M. Foretich, R.M. Bjelland, C.M.F. Durif & C.J. O’Brien. Ontogeny of orientation in Atlantic haddock (Melanogrammus aeglefinus) larvae measured in situ.

Effect of petrochemicals on swimming and orientation of Atlantic haddock (Melanogrammus aeglefinus) larvae measured in situ.

Paris, C.B., H.I. Browman, A.B. Skiftesvik, S. Shema, M. Foretich, R.M. Bjelland, C.M.F. Durif & C.J. O’Brien & S. Meier. Effect of petrochemicals on swimming and orientation of Atlantic haddock (Melanogrammus aeglefinus) larvae measured in situ.

Effect of seismic air gun shooting on Calanus finmarchicus.

Opstad, I., D.M. Fields, H.I. BROWMAN, A.B. Skiftesvik, & S. Shema. Effect of seismic air gun shooting on Calanus finmarchicus.

Host recognition mechanisms in the sea lice: genomic and functional approaches to identify novel chemoattractants derived from salmon skin.

Núñez-Acuña, G., D.M. Fields, S. Shema, H.I. BROWMAN, A.B. Skiftesvik & C. Gallardo-Escárate. Host recognition mechanisms in the sea lice: genomic and functional approaches to identify novel chemoattractants derived from salmon skin.

The structural integrity of the exoskeleton of an ecologically critical planktonic species in North Atlantic ecosystems, Calanus finmarchicus, is unaffected by high CO2.or temperature.

Mount, A., V. Chan, J.A: Runge, D.M. Fields, C. Thompson, S. Shema, R.M. Bjelland, C.M.F. Durif, A.B. Skiftesvik & H.I. Browman. The structural integrity of the exoskeleton of an ecologically critical planktonic species in North Atlantic ecosystems, Calanus finmarchicus, is unaffected by high CO2.or temperature.

Temperature effects on metabolic rate of Calanus spp.

Fields, D.M., H.I. BROWMAN, A.B. Skiftesvik & S. Shema. Temperature effects on metabolic rate of Calanus spp.

Effect of ocean acidification on the grazing rates of Calanus spp. feeding on lithed and delithed coccolithophores.

Fields, D.M., H.I. BROWMAN, A.B. Skiftesvik & S. Shema. Effect of ocean acidification on the grazing rates of Calanus spp. feeding on lithed and delithed coccolithophores.

Magnetic orientation in juvenile European lobster, Homarus gammarus.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik, A.-L. Agnalt & H.I. BROWMAN. Magnetic orientation in juvenile European lobster, Homarus gammarus.

The magnetic compass of glass eel (Anguilla anguilla) is entrained to and modulated by the tidal cycle.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik & H.I. BROWMAN. The magnetic compass of glass eel (Anguilla anguilla) is entrained to and modulated by the tidal cycle.

Orientation of European glass eels (Anguilla Anguilla) arriving at the Norwegian coast.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik & H.I. Browman. Orientation of European glass eels (Anguilla Anguilla) arriving at the Norwegian coast.

Magnetic orientation in juvenile lumpfish, Cyclopterus lumpus.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik & H.I. BROWMAN. Magnetic orientation in juvenile lumpfish, Cyclopterus lumpus.

Glass eels (Anguilla anguilla) orientation is correlated with the position of the moon, even when they cannot see it.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik & H.I. Browman. Glass eels (Anguilla anguilla) orientation is correlated with the position of the moon, even when they cannot see it.

The proteome of Calanus finmarchicus is unaffected by elevated pCO2.

Dineshram, R., V. Thiyagarajan, D.M. Fields, J.A. Runge, C. Thompson, S. Shema, R.M. Bjelland, C.M.F. Durif, A.B. Skiftesvik & H.I. BROWMAN. The proteome of Calanus finmarchicus is unaffected by elevated pCO2.

The effects of an anti-sea lice chemotherapeutant, hydrogen peroxide, on mortality, escape response and oxygen consumption of Calanus spp. Environmental Science and Technology.

Escobar, R.H., D.M. Fields, H.I. Browman, S.D. Shema, R.M. Bjelland, A.B. SKIFTESVIK & C.M.F. Durif. The effects of an anti-sea lice chemotherapeutant, hydrogen peroxide, on mortality, escape response and oxygen consumption of Calanus spp. Environmental Science and Technology.

Learning in juvenile European lobster, Homarus gammarus., is negatively affected by exposure to hydrogen peroxide.

Cresci, A, C.B. Paris, C.M.F. Durif, S. Shema, R. Bjelland, A.B. Skiftesvik, A.-L. Agnalt, O. Samuelsen & H.I. Browman. Learning in juvenile European lobster, Homarus gammarus., is negatively affected by exposure to hydrogen peroxide.

The effects of an anti-sea lice chemotherapeutant, hydrogen peroxide, on mortality, escape response and oxygen consumption of Calanus spp.

Escobar, R.H., D.M. Fields, H.I. Browman, S.D. Shema, R.M. Bjelland, A.B. Skiftesvik & C.M.F. Durif. The effects of an anti-sea lice chemotherapeutant, hydrogen peroxide, on mortality, escape response and oxygen consumption of Calanus spp. Environmental Science and Technology (in preparation).

Last updated: July 18, 2017